Hechtia atlanticasp. nov. (Bromeliaceae), a new species from Veracruz, Mexico
Hechtia atlantica Hern.-Cárdenas, Espejo
& López-Ferr., sp. nov., a new taxon from the state of Veracruz in
Mexico, is described and illustrated. The proposed species is compared
with H.myriantha Mez and H. schottii Baker, taxa with some
similarities. Hechtiaatlantica sp. nov. differs
from these by the size of the rosette and leaf sheath, the shape of
the sepals and petals of the staminate flowers, and the flowering
season. A morphological description and images of the new species are
provided, as well as a comparison table, an identification key, a list
of examined specimens, and a distribution map of all Hechtia Klotzsch species present in
Veracruz.
KEYWORDS: Bromeliaceae, endemism, Mexico, monocots, Poales, Veracruzan Province, new species.
Hechtiaatlantica Hern.-Cárdenas, Espejo &
López-Ferr., sp. nov., un nouveau taxon de l’État de Veracruz au
Mexique, est décrit et illustré. L’espèce proposée est comparée à H. myriantha Mez et H. schottii Baker, taxons présentant quelques
similitudes. Hechtiaatlantica sp. nov. s’en
distingue par la taille des rosettes, de la gaine, la forme des
sépales et des pétales des fleurs staminées ainsi que par la saison de
floraison. Une description morphologique et des images de la nouvelle
espèce sont fournies, ainsi qu’un tableau comparatif, une clé
d’identification, une liste des spécimens examinés et une carte de
répartition de toutes les espèces d’Hechtia Klotzsch présentes dans l’État de
Veracruz.
MOTS CLÉS: Bromeliaceae endémisme, Mexique, monocotylédones, Poales, province de Veracruz, nouvelle espèce.
et al.
2007)et al.
2025), 95 (98 %) of which geographically restricted to
Mexico (updated from Espejo-Serna 2012), and 45 are known only from one
Mexican state. The states with the highest number of et
al. 2020).
In fact, many et
al. 2017a, b). In the last 18 years (2007-2025), knowledge
regarding the systematics and diversity of the genus has increased
significantly, with more than 50 species being described, 47.6 % of
taxa known to date (Espejo-Serna et
al. 2007, 2008; Ramírez-Morillo 2008; López-Ferrari et al. 2009; Martínez-Correa et al. 2010; Burt-Utley et al. 2011; Ramírez-Morillo et al. 2011, 2013, 2014, 2015, 2016, 2021, 2023a, b, 2024; Burt-Utley 2012; Ramírez-Morillo & Jiménez
Nah 2012; López-Ferrari & Espejo-Serna 2013, 2014; García-Ruiz et al. 2014; González-Rocha et al. 2014; Flores-Argüelles et al. 2019, 2024; Hernández-Cárdenas et al. 2019, 2020, 2022, 2023, 2024a, b) (Fig. 1).
For the state of Veracruz, five et al. 2005; Espejo-Serna &
López-Ferrari 2011). As a result of botanical explorations in
Veracruz to prepare the manuscript of the et
al. 2005), we collected, in 2004, at la Quebrada,
municipality of Actopan, specimens of the taxon here proposed as new
(A.Espejo et al. 6726♀; CICY, IEB, UAMIZ, XAL); and although
since then we had suspected that the material could belong to an
undescribed taxon, we did not have, at that time, fertile specimens
of both sexes so we decided to provisionally identify it as
Staminate and pistillate plants of the new
taxon were collected in the municipality of Actopan, Veracruz. The
specimens were processed, measured, analyzed, and a complete
description was made from dry material. Terminology used is that
proposed by Radford et al. (1974) and
Scharf & Gouda
(2008). The types were deposited at UAMIZ and XAL herbaria
(acronyms follow Thiers
[2025]). To ensure the identity of the new species we revised
herbarium material of
Protologues, live and herbarium specimens,
including type material, of all et al. 2005; Espejo-Serna &
López-Ferrari 2011, 2018; Espejo-Serna et
al. 2020): et al. 2007]),
BM The Natural History Museum, London;
C Natural History Museum of Denmark, Copenhagen;
CH El Colegio de la Frontera Sur, San Cristóbal de las Casas;
CHAPA El Herbario-Hortorio del Colegio de Postgraduados, Mexico;
CHIP Instituto de Historia Natural, Tuxtla Gutiérrez;
CIB Universidad Veracruzana, Mexico, Veracruz, Xalapa;
CICY Centro de Investigación Científica de Yucatán, Mérida;
CITRO Centro de Investigaciones Tropicales, Universidad Veracruzana, Xalapa;
CORU Universidad Veracruzana, Campus Córdoba, Córdoba;
F Field Museum of Natural History, Chicago;
GH Harvard University, Cambridge;
IBUG Universidad de Guadalajara, Zapopan;
IEB Instituto de Ecología, Pátzcuaro;
K Royal Botanic Gardens, Kew;
MEXU Universidad Nacional Autónoma de Mexico, Mexico;
MO Missouri Botanical Garden, Saint Louis;
MICH University of Michigan, Ann Arbor;
NY The New York Botanical Garden, New York;
SEL Marie Selby Botanical Gardens, Sarasota;
TEX University of Texas at Austin, Austin;
UADY Universidad Autónoma de Yucatán, Mérida;
UAMIZ Universidad Autónoma Metropolitana Iztapalapa, Mexico;
UC University of California, Berkeley;
US Smithsonian Institution, Washington;
WIS University of Wisconsin-Madison, Madison;
XAL Instituto de Ecología, Xalapa.
Artificial key to the
- 1. Leaf margin minutely serrate; peduncle 3-5 mm diameter,
inflorescence pendulous ⇒
- — Leaf margin conspicuously spinose; peduncle 8-30 mm diameter, inflorescence erect ⇒ 2
- 2. Primary branches glomerate to very slightly elongate, 1.5-6.5 cm long ⇒ 3
- — Primary branches never glomerate, 10-40 cm long ⇒ 4
- 3. Leaf blades 15-31 cm long; pedicels glabrous, petals 6-7
mm long ⇒
- — Leaf blades 37-60 cm long; pedicels lepidote, petals 3.5-5
mm long ⇒
- 4. Peduncle, floral bracts, sepals, and petals glabrous ⇒
- — Peduncle, floral bracts, sepals, and petals lepidote ⇒ 5
- 5. Leaf blades 100-170 cm long; inflorescence twice to three
times branched; floral bracts 3-5 mm long,
c.3 mm wide ⇒ - — Leaf blades 50-60 cm long; inflorescence twice branched,
never thrice; floral bracts 5.5-6.5 mm long, 4.5-5 mm wide ⇒
-
-
The specific epithet refers to the region where the new taxon inhabits, located at the proximity of the Atlantic Ocean, in the coast of Gulf of Mexico.
2978 ♀;
UAMIZ•
sensuRzedowski
1978). Plants of the new species grow in colonies or
sometimes solitary on the sidewalls along the edge of la
Quebrada, between 0 and 35 m elevation.
Plants saxicolous to terrestrial, in
flower 150-250 cm high, rosettes acaulescent, 50-60 cm high,
40-70 cm diam., cespitose forming clumps of five to ten
rosettes or sometimes solitary. Leaves 40 to 60, ascending,
recurved toward the apex; sheaths yellowish-brown at the
base to brown toward the distal part, very broadly ovate,
10-13 cm long, 12-16 cm wide, glabrous near the base and
lepidote abaxially toward the apex, glabrous adaxially, with
minute marginal sharp teeth; blades grayish-green, narrowly
triangular, long attenuate, 50-60 cm long, 8.5-10 cm wide at
the base, very densely white lepidote abaxially, glabrous
adaxially, margins with divaricate to ascending green to
brown sharp teeth, 5-10 mm long, 5-8 mm wide at the base,
1-1.5 cm apart. Inflorescences lateral, erect, twice
branched in staminate and pistillate plants. Staminate
inflorescence 170-190 cm long; peduncle brown, terete, 1.3-1.8
cm diam., lepidote, internodes 2-6 cm long; pedunclebracts brown,
foliaceous, sheaths triangular, 2-5 cm long, 1.4-2.2 cm
wide, lepidote on both surfaces, margins hyaline, blades
narrowly triangular, 1.5-3.5 cm long, 0.7-1.5 cm wide,
lepidote on both surfaces, entire, the basal ones longer
than the internodes, the upper ones shorter than the
internodes; primarybracts light brown, narrowly
triangular, 1-3.2 cm long, 0.7-1 cm wide, caudate, entire,
margins hyaline, lepidote on both surfaces; main axis 110-140
cm long, internodes 2-6 cm long; primarybranches 25 to 30, 15-25 cm long,
1-1.2 cm diam., secondary branches 1 to 7, 5-9 cm
long, 0.8-1 cm diam.; floralbracts brown, broadly ovate, c. 5.7 mm long, c. 4.5 mm wide,
longer than pedicels, apiculate, entire to erose, lepidote
on both surfaces. Staminate flowers 20 to 40 per primary
branch, sessile to shortly pedicellate, pedicelsc. 0.5 mm long, inconspicuous;
sepals brown,
ovate, c. 5 mm
long, c. 4 mm
wide, apiculate to acute, hyaline, entire, lepidote on both
surfaces; petals white, broadly elliptic,
4.7-5 mm long, 4-4.5 mm wide, obtuse, entire, glabrous
adaxially, lepidote abaxially; stamens equal in length; filaments white,
narrowly triangular, flattened, 3.5-4 mm long; anthers green,
oblong, 2-2.3 mm long, versatile, pollen yellow; pistillode
conspicuous, white, lepidote. Pistillate inflorescence
100-190 cm long; peduncle brown, terete, 0.7-1 cm
diam., lepidote, internodes 1.5-2.5 cm long; pedunclebracts brown,
foliaceous, sheaths triangular, 2-3 cm long, 1-1.5 cm wide,
lepidote on both surfaces, margins hyaline, blades linear,
1-1.5 cm long, 0.5-1 cm wide, lepidote on both surfaces,
entire, the basal ones longer than the internodes, the upper
ones shorter; primarybracts brown, triangular, 1-2 cm
long, 0.5-1 cm wide, caudate, margins entire, hyaline,
lepidote on both surfaces; main axis 50-60 cm long,
internodes 2.5-3.5 cm long; primarybranches 12 to 18, 8-25 cm long,
0.8-1.2 cm diam.; secondary branches 1 to 6, 2-9 cm
long, 0.8-1 cm diam.; floralbracts brown, broadly ovate, c. 6 mm long, c. 5.3 mm wide,
longer than pedicels, apiculate, lepidote on both surfaces,
margins hyaline, entire to erose. Pistillate flowers 20 to
30 per primary branches, sessile to shortly pedicellate, pedicelsc. 0.5 mm long; sepals brown,
ovate, 5.5-7 mm long, 3.5-4 mm wide, acute, lepidote on both
surfaces, margins hyaline, entire; petals white, broadly ovate,
6-6.4 mm long, c.
4.7 mm wide, acute, entire, glabrous adaxially,
lepidote abaxially;
staminodes rudimentary, white; ovary superior, green to white,
oblong, c. 4.5
mm long, c. 2.8
mm diam., lepidote;
stylar branches white, recurved, slender, 1.2-1.5 mm
long, stigmas conduplicate-patent, papillose. Capsules dark
brown, ovoid, 7-7.5 mm long, 4-5 mm diam.; seeds not seen.
The specimens of the new taxon never
exceed in flower 2.5 m height, they are cespitose and have
acaulescent rosettes that produce lateral shoots. The
rosettes are less than 60 cm high and 70 cm in diameter,
being much smaller than those of sensuRzedowski 1978)
facing the sea and blooms from July to August.
On the other hand, the plants of et al. 2020) (Figs 7; 8). It grows in tropical
deciduous forests (sensuRzedowski
1978).
et al. 2005,
2020). c. 100 cm) of the rosette, in the
larger leaf sheath (10-13 × 12-16 vs 4-5 × 4-5 cm), in the
wider leaf blade (8.5-10 vs 3.4-4.5 cm), in the shape
(widely elliptic vs obovate) and lepidote (vs glabrous)
staminate petals, in the lepidote pistillate petals (vs
glabrous), and in its geographical distribution (Veracruz vs
Campeche and Yucatán) and flowering season (July to August
vs April to May) (Table
1).
To the curators of the herbaria BM, C, CH, CHIP, CIB, CICY, CITRO, CORU, F, GH, IBUG, IEB, K, MEXU, MO, MICH, NY, SEL, TEX, UADY, UAMIZ, UC, US, WIS, and XAL, for allowing us access to their collections. To María Teresa Jiménez Segura for the drawings of figures 4 and 8. To Elton M. Carvalho Leme and Walter Till for their observations and suggestions that improved the manuscript.
Mexico • Veracruz, Acultzingo; c. 5
km después de Acultzingo rumbo a Tehuacán; 18°42’55”N, 97°20’12”W; 2035 m alt.;
26.IV.2004; fr; A.
Espejo-Serna et al. 6746♀; UAMIZ57697•Acultzingo, Km
331.4 de la carretera Tehuacán-Orizaba. 4 km después de
Puerto del Aire; 18°42’53”N, 97°20’11”W; 2013 m alt.;
10.XII.2005; fr; N.
Martínez et al. 22♀; UAMIZ66421, XAL138898•
Acultzingo, cerro por el camino a El Potrero, 18°42’10”N, 97°16’17”W; 1887 m alt.;
1.IX.2012; fr.; J.
Rivera Hernández & A. Vargas 4848♀; CORU, UAMIZ 74217• Acultzingo,
about 2.4 miles west end of Acultzingo along MEX 150 which
enters Puebla and junction with MEX 125 to Tehuacán; 18°42’50.2”N, 97°20’5.3”W; 2000 m alt.;
4.VIII.1991; J.
Utely & K. Utley 8748♀; MO6881026.
Mexico • Puebla, Chignautla, bei
Chignautla; 19°48’52”N, 97°23’17”W; 7000-8000 ft;
VII.1841; F.
Liebmann [s.n.]; holotype: C; isotype: B10
0242805•Guadalupe Victoria, a 6.6 km al SW de Alchichica, c.
2 km al SW de la desviación a Techachalco, carretera San
Salvador el Seco – Perote; 19°22’10”N, 97°26’15”W; 2390 m alt.;
22.IV.2004; A.R.
López-Ferrari et al. 3109♀; UAMIZ57764•Veracruz, Perote,
carretera que va de Amozoc a Xalapa (140D); 19°32’12.4”N, 97°22’39.6”W; 2414 m alt.;
4.XII.2022; R.
Hernández-Cárdenas & S. Lara-Godínez 2629♀; fr; UAMIZ88992•
Perote, Tenextepec, cerro de Tenextepec; 19°28’25.8”N, 97°16’49.8”W; 2550 m alt.;
3.III.1973; G.
Castillo-Campos et al. 14257♀; XAL[s.n.]•Perote, hacienda de San
Agustín, al SE; 19°27’47”N, 97°17’10”W; 2560 m; 4.XI.1998;
fr.; G.
Castillo-Campos et al. 18732♀; MEXU[1359682]•Perote, cerros
calizos cerca del límite con el estado de Puebla (cerca de
Alchichica); 19°20’29.6”N, 97°28’8”W; 2430 m alt.;
14.VII.1968; C.H.
Ramos 227♂; MEXU307849.
Mexico • Veracruz, Actopan, cerro de
la Mesa; 19°32’0”N, 96°30’0”W; 400 m alt.;
5.VI.1987; R. Acosta
P. 1607♂;
fl; CIB, XAL[s.n.]•Apazapan, 0.5 km de
Apazapan hacia Jalcomulco; 19°19’8.3”N, 96°44’17”W; 378 m alt.;
24.IV.2007; I.
Ramírez et al. 1462♀; CHIP, XAL105318•Camarón de
Tejeda, Camarón; 19°1’23”N, 96°36’52”W; 320 m alt.;
1.III.1921; C. A.
Purpus 8628♂; fl.; GH[s.n.], MO, UC•Emiliano Zapata,
cañón de Palo Gacho; 19°23’3”N, 96°37’30”W; 400 m alt.;
1.III.1981; fl., M.
Cházaro & A. Justo 1439♂; XAL82714•Emiliano Zapata,
Cerro Gordo, carretera Jalapa – Veracruz; 19°31’54.5”N, 96°39’32.25”W; 420 m alt.;
24.I.1972; fl.; J.
Dorantes 499♂; MEXU172424•Emiliano Zapata, La
Cumbre, carretera Xalapa – Veracruz; 19°23’11”N, 96°39’3”W; 374 m alt.;
5.III.2004; fl.; A.
Espejo-Serna & J. García-Cruz 6704♀; IEB, MEXU1351036, MEXU1351037,
MEXU1351038, MEXU1351092, UAMIZ57749, XAL•ibid.6705♂; IEB, MEXU1351073, MEXU1351075,
MEXU1351094, MEXU1351094, UAMIZ57754, XAL•Emiliano Zapata,
Carrizal; 19°21’50”N, 96°38’5”W; 420 m alt.;
12-14.V.1901; fl.; E. A. Goldman 712♂ F,
US397082•Emiliano Zapata, between Plan del Río and La
Cumbre, on Xalapa-Veracruz road; 19°23’35.8”N, 96°39’4.2”W; 340 m alt.;
14.I.1993; fl.; P.
Hietz & U. Seifert 815♂; XAL82716•Emiliano Zapata,
Cerro Gordo, carretera Xalapa-Veracruz, entre Cerro Gordo y
Plan del Río; 19°26’0”N, 96°41’53”W; 420 m alt.;
2.III.1998; I.
Ramírez & C. Durán 642♂; XAL105317•ibid.643♀; XAL147240•Emiliano Zapata,
Cerro Gordo; 19°26’0”N, 96°41’53”W; 450 m alt.;
15.I.1975; fl.; F.
Ventura A. 10803♂; IEB, MEXU349860, XAL•
Jalcomulco, cerro de Achichuca, Tuzamapam; 19°23’N, 96°48’5”W; 800 m alt.;
2.III.1979; fl., fr.; G. Castillo-Campos 463♀/♂; CIB, US2854481, US2854482,
XAL82713•Jalcomulco, 4 km antes de Tacotalpan, camino a
Jalcomulco; 19°21’25.9”N, 96°46’17”W; 6345 m;
9.XI.1991; G.
Castillo C. & P. Zamora C. 7591; XAL[s.n.]•Jalcomulco,
al N del río de los Pescados, frente al puente; 19°21’53.2”N, 96°49’32.7”W; 460 m alt.;
18.VI.1998; fr; G.
Castillo-Campos et al. 17918♂; CICY, IEB159178, XAL109083•
Jalcomulco, subida al cerro de Achichuca, al NW de
Jalcomulco; 19°21’4”N, 96°46’45”W; 800 m alt.;
6.III.2004; fl.; A.
Espejo-Serna et al. 6709♀; UAMIZ567730•Jalcomulco,
barranca de Santa María Tlatetla prope Mirador; 19°16’54”N, 96°43’30”W; III.1842; F. Liebmanns. n.; B10 0242806,
C•Jalcomulco, camino a Coetzalan-Cueva del Abono; 19°18’0”N, 96°42’30”W; 500 m alt.;
12.II.1983; fl.; L.
Robles H. 2♂; XAL82722•Jalcomulco, cerro de
Achichuca, ejido Tuzamapan, sobre carretera Tuzamapan-
Huatusco, antes del río Pescado; 19°22’38”N, 96°49’12”W; 600 m alt.;
9.V.1981; fr.; A. P.
Vovides y J. I. Calzada 657♀; XAL82704•Puente Nacional,
parte alta de la barranca del Pastor, c. 2.5 km después de
Mata de Jobo, rumbo a Santa María Tatetla; 19°15’6”N, 96°41’26”W; 500 m; 22.IV.2004;
fr.; J. Ceja-Romero
et al. 1636♂; IEB, MEXU1355924, MEXU1356096,
UAMIZ57706, XAL•ibid.1637♀; IEB, UAMIZ57729•Jalcomulco,
carretera federal que va de Jalcomulco a Apazapan; 19°20’06”N, 96°45’06”W; 350 m alt.;
8.II.2025; fl; R.
Hernández-Cárdenas & S. Lara-Godínez 3210♂; XAL, ibid. 3211♀; XAL•Puente
Nacional, camino de herradura entre el rancho Dos Caminos y
la barranca de Panoaya; 19°12’7”N, 96°40’25”W; 790 m; 25.IV.2004;
fl.; A.R.
López-Ferrari et al. 3119♀; MEXU1355921, UAMIZ57707•
Puente Nacional, barranca de Panoaya, 1.5 km al NW de El
Coyol; 19°12’7”N, 96°40’25”W; 420 m alt.;
30.I.1986; M.E. ♂; CORU, XAL•Puente Nacional,
barranca near La Palmilla; 19°11’57.5”N, 96°45’4”W; 590 m alt.;
1.III.1934; fl.; C.A. Purpus 16359 GH[s.n.]•Totutla,
Fortín; 19°12’41”N, 96°51’2”W; 940 m alt.;
III.1919; fl.; C.A.
Purpus8505♂; GH, MO, UC204964•Totutla,
Zacuapan; 19°12’36.7”N, 96°50’38.9”W; 915 m alt.;
I.1920; fl.; C.A.
Purpus 8522a♂; MO858645.
Mexico • Veracruz, Jalcomulco, cerro
de Achichuca, entre Jalcomulco y Tuzamapan; 19°23’0”N, 96°48’40”W; 600 m alt.;
17.IV.1984; fr.; G.
Castillo-Campos 3124♀; CIB, XAL[s.n.]•Jalcomulco, ejido de
Jalcomulco, 100 m antes de llegar a La Mesa del Barro; 19°21’55”N, 96°46’30”W, 600 m alt.;
18.IX.1980; fr.; G.
Castillo-Campos & L. Tapia 1141♀; XAL[s.n.]•Jalcomulco, 1.5 km al NO
del poblado de Jalcomulco; 19°20’47”N, 96°46’37”W; 600 m alt.;
17.VI.1998; fl.; G.
Castillo-Campos et al. 17881♀; IEB, MEXU, UAMIZ54803,
XAL109082•Jalcomulco, 5 km al S de Santa María Tatetla; 19°15’3”N, 96°41’27”W; 450 m; 3.IX.1998;
fr.; G.
Castillo-Campos et al. 18247♀; IEB, MEXU1085949, XAL109081•
Jalcomulco, Santa María Tatetla – Coetzalan; 19°17’50”N, 96°42’30”W; 450 m alt.;
12.IV.1983; fl.; L.
Robles H. 85♂; IBUG92013, XAL[s.n.]•Jalcomulco, 4
km antes de Jalcomulco viniendo de Apazapan; 19°19’34.9”N, 96°43’36.2”W; 306 m alt.;
III.2011; fl.; I.
Ramírez & G. Carnevali 1667♂; MEXU, XAL147241•Puente
Nacional, c. 8.5 km después de Mata de Jobo, rumbo a Santa
María Tatetla; 19°15’16”N, 96°41’19”W; 467 m alt.;
22.IV.2004; fl.; J.
Ceja et al. 1642♂; UAMIZ57766•ibid. fr.; 1643♀; IEB, MEXU, UAMIZ57767•Puente
Nacional, barranca de Pachuquilla; 19°14’0”N, 96°36’40”W; 450 m alt.;
14.IV.1990; fl.; M.
Cházaro & C. Viveros 6211♂; MEXU565358, MICH, TEX, WIS,
XAL[s.n.]•
Puente Nacional, barranca de Panoaya; 19°12’7”N, 96°40’25”W; 795 m alt.; 25.IV
2004; fl.; A.R.
López-Ferrari 3120♀; UAMIZ57768•Puente Nacional,
barranca de Panoaya, 1.5 km al NW de El Coyol; 19°11’9.34”N, 96°41’26”W; 476 m alt.;
30.I.1986; fr; M.E. s.n.]•Tenampa, barranca de
Tenampa; 19°14’15”N, 96°52’50”W; 990 m alt.;
1.V.1919; fl.; C.A.
Purpus 8420♀; GH, MO, NY247109, UC, US•
Tlacotepec de Mejía, barranca de Consoquitla, prope el
Fortín; 19°12’5.54”N, 96°50’13.4”W; 830 m alt.;
III.1842; F.
Liebmann 7951; C, F.
Mexico • Campeche, Calakmul, 1 km del
ejido Eugenio E. Castello – 2; 18°41’36”N, 89°35’19”W; 248 m alt.;
25.V.2003; fl.; D.
Álvarez 5076♂; MEXU1237845, UAMIZ56294•
Calakmul, 3 km al W de Eugenio Echeverría Castellot I, sobre
la carretera Escárcega – Chetumal; 28.X.1997; fl; E. Martinez
29610♀/♂; CHAPA, CHIP23612, MEXU908973,
MEXU909129, MO5704619•Calakmul, mina de yeso Coconal, a 23
km al W de Xpujil, camino a Escárcega; 18°31’47”N, 89°35’21”W; 262 m alt.;
12.VIII.1997; fl.; E. Martínez 28433♂; CHAPA, CICY,
MEXU837905•Yucatán, Maxcanú, 20°34’36.9”N, 90°0’48.1”W; 60 m alt.; 1917;
fl.; G.F. Gaumer
& sons 23839; GH[s.n.]•Maxcanú, cerro en el sur
del pueblo, capilla la Ermita; 20°34’33”N, 90°0’16”W; 51 m alt.;
10.IV.2011; I.
Ramírez et al. 1676♂; CICY, IEB, MEXU, MO, US, XAL•
ibid. 1677♀ CICY•Maxcanú,
cerro de Maxcanú; 20°34’15.3”N, 90°1’25.6”W; 55 m alt.;
1865; A.C. V. Schott
645; BM85619•Opichén, grutas de Calcehtok; 20°33’36.7”N, 89°56’27”W; 100 m alt.;
10.XI.1980; fr.; J.I. Calzada et al. 6712♀; CICY, XAL•
Opichén, Okintok; 20°33’32.5”N, 89°57’13.9”W; 43 m alt.; S.P. Darwin & E.
Sundell 2068; MEXU, MO3032530, MO3032531•Opichén,
cultivated at Mérida; 14.II.2004; fl; A. Espejo & A.R. López-Ferrari
6701♀; UAMIZ•6702♂; UAMIZ•Opichén,
grutas de Calcehtok; 20°33’36.7”N, 89°56’27”W; 100 m alt.;
24.II.1991; fl.; M.
Méndez et al. 471♂; TEX46072, XAL[s.n.]•Opichén,
grutas de Calcethock; 20°33’N, 89°48’W; 100 m; 17.III.1984; fr.;
J.J. Ortiz 437;
MEXU650753, MO.
Mexico • Veracruz, Actopan, cerro de
la Mesa (sierra Manuel Diaz); 19°32’0”N, 96°30’0”W; 400 m alt.;
6.VI.1987; fl.; A.
Acosta P. 1599♂; XAL•Actopan, raya La Tempestad
(sierra Manuel Diaz); 19°32’0”N, 96°27’0”W; 400 m alt.;
22.XII.1985; fr.; A.
Acosta & N. Acosta 140♀; UAGC8255, XAL[s.n.]•Actopan,
trapiche del Rosario; 19°33’23”N, 96°44’28”W; 450 m alt.;
31.V.2000; fl; G.
Castillo-Campos 19906♂; XAL109091•Actopan, 3 km aprox.
al NE de la desviación a Otates; 19°32’53”N, 96°42’33”W; 400 m alt.;
30.IX.1999; fr.; G.
Castillo-Campos & I. Acosta R.19397♀; MEXU, XAL[s.n.]•Actopan,
trapiche del Rosario; 19°32’45”N, 96°44’5”W; 440 m alt.;
22.X.1999; fr.; G.
Castillo-Campos & I. Acosta R 19539♀; XAL109086,
XAL109087•ibid.19580♀; XAL109084, XAL109085• Actopan,
Sierra Manuel Díaz; 19°32’36”N, 96°26’17”W; 200 m alt.;
2.IV.1998; fl.; G.
Castillo-Campos et al. 17644♀; CICY, XAL[s.n.]•Actopan, el Común en las
faldas de la sierra Manuel Díaz, entre Mozomboa y la
Costera; 19°31’3.6”N, 96°27’19.6”W; 100 m alt.;
12.X.1984; fr.; G.
Castillo-Campos et al. 3403♀; CIB, UAGC8260, XAL[s.n.]•Actopan,
sierra de Manuel Díaz; 19°32’36”N, 96°26’17”W; 200 m alt.;
1.IV.1998; fr.; G.
Castillo-Campos et al. 17590♀; CICY, XAL[s.n.]•Actopan, trapiche del
Rosario;•Actopan, 19°32’21.5”N, 96°44’38.8”W; 530 m alt.;
14.V.2004; fl; G.
Castillo-Campos & B. Cano E. 20916♂; UAMIZ58436•ibid.20918♀; UAMIZ58439•ibid. 20919♂; UAMIZ58437•Actopan,
alrededores de Trapiche del Rosario; 19°32’47”N, 96°44’8”W; 495 m alt.;
23.IV.2004; fl.; A.
Espejo-Serna et al. 6731♀; MEXU1350553, MEXU1350570,
UAMIZ57759, XAL •
ibid.6734♂; UAMIZ57755•Emiliano Zapata,
lomas de Rogel, al N; 19°33’34”N, 96°47’37”W; 640 m alt.;
20.VI.1999; fr.; G.
Castillo-Campos et al. 19217♀; XAL109089, XAL109090• Emiliano
Zapata, parte alta de la cascada de Cerro Gordo (río San
Antonio). Aproximadamente 2 km al NE de Cerro Gordo; 19°26’55”N, 96°41’13”W; 420 m alt.;
4.XI.2007; fr.; P.
Carrillo-Reyes & D. Cabrera-Toledo 5242♀; UAMIZ69335,
XAL91492•Emiliano Zapata, los Reyes; 19°27’5”N, 96°43’54”W; 670 m; 24.VII.1971;
fr.; F. Ventura A.
3946-B♀;
ENCB[s.n.], IEB
•Jalcomulco, 2 km al N de Jalcomulco; 19°20’48”N, 96°46’19.5”W; 435 m alt.;
2.IX.1998; fr.; G.
Castillo-Campos et al. 18179♀; XAL[s.n.]•Naolinco, orillas de la
carretera Actopan km 21 antes de la desviación al Coyol;
19°35’21”N, 96°50’20”W; 530 m alt.;
26.VI.1976; fr.; R.
Ortega O. 306♀; ENCB, MEXU, MO3504273, XAL•
Puente Nacional, Tamarindo; 19°20’21”N, 96°29’25”W; 144 m alt.;
20.VI.1981; fl.; A.
Figueroa N. et al. 23♂; XAL82719•Puente Nacional, 1 km
antes Encinal, carretera Puente Nacional – Huatusco; 19°12’30”N, 96°48’55”W; 700 m alt.;
5.VII.1971; fr.; L.I. Nevling & A. Gómez-Pompa
2405♀;
MEXU166404•Vega de Alatorre, 2 km antes de La Mesilla,
camino Los Reyes-Santa Gertrudis; 19°50’30”N, 96°33’30”W; 350 m alt.;
24.VII.1981; fr.; A.
Benavides & G. Castillo-Campos 83♀; CORU, XAL[s.n.]•Xalapa, malpays de
Chiltoyac « Chiltoyaque »; 19°34’46.7”N, 96°52’8.5”W; 1030 m alt.;
IV.1929; fl.; C.
Schiede & F. Deppe s.n.; B10 0144793, HAL.
A-
C, habit;
D,
E, plants at type locality. Scale bar: 20 cm. Photo credits: R. Hernández-Cárdenas.
A, rosette with lateral inflorescence;
B, pistillate inflorescence;
C, pistillate flowers;
D, staminate flowers. Scale bars: A, 10 cm; C, D, 1 cm. Photo credits: R. Hernández-Cárdenas.
:
A, staminate and pistillate plants in bloom;
B, pistillate primary branch;
B1, pistillate flower;
B2, floral bract;
B3, sepals;
B4, petals;
B5, pistil;
C, staminate primary branch;
C1, staminate flower;
C2, floral bract;
C3, sepals;
C4, petals;
C5, stamens. Scale bars: A, 20 cm; B, C, 5 cm; B1-B5, 7 mm; C1-C5, 6 mm. Illustration drawing A, B, B1, C, C1 by M. T. Jiménez Segura; photo credits C2-C5, B2-B5 by R. Hernández-Cárdenas.
A,
B,
A, plant in the habitat;
B, staminate flowers;
C, pistillate flowers. Scale bars: A, 100 cm; B, C, 5 mm. Photo credits: R. Hernández-Cárdenas.
A, staminate and pistillate plants in bloom;
B, staminate primary branch;
C, pistillate primary branch;
D, staminate flower;
D1, floral bract;
D2, sepals;
D3, petals;
D4, stamens;
E, pistillate flower;
E1, floral bract;
E2, sepals;
E3, petals;
E4, pistil. Scale bars: A, 1 m; B, C, 10 cm; D, E, 4 mm; D1-E4, 3 mm. Illustration drawing by M. T. Jiménez Segura.
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Hechtiaspecies from Oaxaca, México.
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Hechtia(Bromeliaceae, Pitcairnioideae). I. New and noteworthy species of
Hechtiafrom Mexico.
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Hechtia(Bromeliaceae; Pitcairnioideae) del centro de México.
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Hechtia aquamarina,
Hechtia caudata,
Hechtia confusa,
Hechtia conzattiana,
Hechtia elliptica,
Hechtia fragilis,
Hechtia galeotti,
Hechtia laevis,
Hechtia lundelliorum,
Hechtia lyman-smithi,
Hechtia melanocarpa,
Hechtia montana,
Hechtia mooreana,
Hechtia myriantha,
Hechtia podantha,
Hechtia pringlei,
Hectia purpusii,
Hechtia reticulata,
Hechtia rosea,
Hechtia sphaeroblasta,
Hechtia stenopetala,
Hechtia subalata. Mapas de distribución potencial. Unidad Iztapalapa, Universidad Autónoma Metropolitana. México Available from: http://www.conabio.gob.mx/informacion/gis [accessed 3 March 2025].
Hechtia aquamarina,
Hechtia caudata,
Hechtia confusa,
Hechtia conzattiana,
Hechtia elliptica,
Hechtia fragilis,
Hechtia galeotti,
Hechtia laevis,
Hechtia lundelliorum,
Hechtia lyman-smithi,
Hechtia melanocarpa,
Hechtia montana,
Hechtia mooreana,
Hechtia myriantha,
Hechtia podantha,
Hechtia pringlei,
Hectia purpusii,
Hechtia reticulata,
Hechtia rosea,
Hechtia sphaeroblasta,
Hechtia stenopetala,
Hechtia subalata. Registros de presencia usados para elaborar el mapa distribución potencial. Unidad Iztapalapa, Universidad Autónoma Metropolitana, México. Available from: http://www.conabio.gob.mx/informacion/gis [accessed 3 March 2025].
Hechtia(Bromeliaceae) de México.
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Hechtia(Hechtioideae, Bromeliaceae) from Jalisco, Mexico.
Phytotaxa414: 105-112. https://doi.org/10.11646/phytotaxa.414.2.2
Hechtia(Bromeliaceae; Hechtioideae) from Jalisco, Mexico.
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Hechtia(Bromeliaceae; Hechtioideae) del estado de Morelos, México.
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Hechtia(Bromeliaceae, Hechtioideae) from Oaxaca, Mexico.
Phytotaxa397: 280-290. https://doi.org/10.11646/phytotaxa.397.4.2
Hechtia(Bromeliaceae; Hechtioideae) from Guerrero, Mexico.
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Hechtia(Bromeliaceae; Hechtioideae) from Puebla, Mexico.
Systematic Botany47:190-207. https://doi.org/10.1600/036364422X16442668423518
Hechtia kruseanaand
Hechtia rosamariae(Bromeliaceae; Hechtioideae), two new species from Guerrero, Mexico.
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Hechtia marabascoensis(Hechtioideae; Bromeliaceae), a novelty from Colima and Jalisco, Mexico.
Phytotaxa659: 87-96. https://doi.org/10.11646/phytotaxa.659.1.7.
Hechtia cerrostlatilpae(Hechtioideae; Bromeliaceae), una nueva especie de Morelos, México.
Botanical Sciences102: 1353-1362. https://doi.org/10.17129/botsci.3513
Hechtia, eine neue Gattung der Bromeliaceen.
Allgemeine Gartenzeitung3: 401-403.
Hechtia mapimiana(Bromeliaceae; Hechtioideae), una nueva especie del estado de Durango, México.
Acta Botanica Mexicana102: 89-97. https://doi.org/10.21829/abm102.2013.230
Hechtia rubicunda(Bromeliaceae; Hechtioideae), una nueva especie de Oaxaca, México.
Acta Botanica Mexicana107: 153-164. https://doi.org/10.21829/abm107.2014.205
Hechtia caulescens(Bromeliaceae), a new species from Central Mexico.
Novon19: 197-200. https://doi.org/10.3417/2007078
Hechtia(Bromeliaceae, Hechtioideae) from Mexico.
Systematic Botany35: 745-754. https://doi.org/10.1600/036364410X539835
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Hechtia(Bromeliaceae) from the States of Querétaro and Hidalgo, Mexico.
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Hechtia aquamarina, a new name for
Hechtia pueblensisI. Ramírez and Jiménez.
Phytotaxa48: 33. https://doi.org/10.11646/phytotaxa.48.1.5.
Hechtia(Bromeliaceae) from Chihuahua, Mexico.
Novon21: 362-367. https://doi.org/10.2307/23018450
Hechtia(Bromeliaceae):
Hechtia sanchezii, a new species from San Luis Potosí, Mexico.
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Hechtia(Bromeliaceae) from southwestern Tamaulipas, Mexico.
Phytotaxa112: 3-42. https://doi.org/10.11646/phytotaxa.112.2.1
Hechtia(Bromeliaceae: Hechtioideae) from Hidalgo (Mexico).
Phytotaxa221: 157-165. https://doi.org/10.11646/phytotaxa.221.2.5
Hechtia(Hechtioideae; Bromeliaceae) from Jalisco, Mexico.
Phytotaxa266: 261-270. https://doi.org/10.11646/phytotaxa.266.4.3
Hechtia(Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico
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Hechtia(Bromeliaceae: Hechtioideae).
Phytotaxa178: 116-122. http://dx.doi.org/10.11646/phytotaxa.178.2.3
Hechtia(Bromeliaceae: Hechtioideae) from the Pacific Lowlands in Mexico.
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Hechtia(Bromeliaceae) from the Mexican state of Jalisco.
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